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− | + | G. 3c) Embryo cleaves equally Blastula is slightly flattened along the | |
+ | G. 3c) Embryo cleaves equally Blastula is slightly flattened along the animal-vegetal axis Gastrula is somewhat flattened along animal-vegetal axis, becomes ciliated, and develops an apical tuft in addition to a vegetal invagination (blastopore). Gastrulae may perhaps swim freely in sophisticated stages (Fig. 3e) Paired cephalic discs invaginate (Fig. 4) Earliest look Earliest look (16 ) (8 ) two.five h 3h 15.five h 20.five h 3h 4h 22 h 24 hCephalic discs30 h 42 h-- -- 3 daysCephalic and trunk discs Paired trunk discs invaginate (Fig. 4) Pileus stageLarva develops transient lobes and lappets, the gut curves backward, the paired three days cerebral organ discs invaginate in the gut, and also the proboscis and dorsal rudiment seem (Figs. 3f and 5). Ciliary bands seem as four segments which span each and every transient lobe and lappet (Fig. six) The head and trunk rudiments fuse about the base in the gut (Fig. 8). Ciliary band segments are re-arranged to kind two full transverse ciliary bands Epidermis of trunk rudiment extends over the proboscis, but has not but fused with the epidermis of the head rudiment, leaving a dorsal gap (Fig. 9) The head and trunk rudiments fused to form a total juvenile (Fig. ten). Juvenile erupts from and devours larval physique in catastrophic metamorphosis 4 days 6 days 9 daysTorus stage Hood stage Metamorphosis-- -- 18 daysDash represents missing databands from the anterior and posterior lobes (Fig. 6b), too as anterior and posterior gaps separating the ciliary bands from the two lappets (Fig. 6a). The ciliary bands spanning every single from the transient lappets extend from a tiny pair of epidermal invaginations involving the anterior lobe and every single lappet (Fig. 5a, c and d, Extra file 2 -- Movie two, Added file three -- Film three). These "pits" are inside a position corresponding towards the anterior axils (the development zones) of the pilidium larva, which give rise for the ciliary bands in common pilidia [19]. At this point, the cephalic discs are positioned above the transient anterior lobe, when the trunk discs are beneath the backward curve with the gut, along the posterior side of every single lappet (Fig. 5b-d). In the same time, the unpaired dorsal rudiment types along the inner pilidial epidermis dorsal/apical to the gut, plus the unpaired proboscis rudiment types amongst the cephalic discs (Fig. 5). The origin of those rudiments is uncertain, but possibly, as is hypothesized for any typical pilidium, they may be mesenchymal [10]. At any price, they don't appear to kind by invagination from the larval epidermis. Moreover, two shallow chambers outpocket in the basal portion in the gut (close to the blastopore) among the building cephalic and trunk discs on either side of the mid-sagittal plane. These invaginations of your gut elongate and kind the cerebral organ discs (Fig. 5e3-f ). The larva also starts to develop musculature at this point. Initially, it develops circumferential muscles underlying the ciliary bands of your transitory larval lobes (which later form the anterior ciliary band). As these muscles type, some start to arc towards the posterior on the larva, extending into every single ofthe transitory lappets, instead of encircling the larva (Fig. 7a-b). These extensions type an "arc" of muscle which follows the curve on the lappet (consider two jump ropes, each and every wrapped around opposite sides on the larva, are permitted to droop into the lappet on its respective side). |
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G. 3c) Embryo cleaves equally Blastula is slightly flattened along the G. 3c) Embryo cleaves equally Blastula is slightly flattened along the animal-vegetal axis Gastrula is somewhat flattened along animal-vegetal axis, becomes ciliated, and develops an apical tuft in addition to a vegetal invagination (blastopore). Gastrulae may perhaps swim freely in sophisticated stages (Fig. 3e) Paired cephalic discs invaginate (Fig. 4) Earliest look Earliest look (16 ) (8 ) two.five h 3h 15.five h 20.five h 3h 4h 22 h 24 hCephalic discs30 h 42 h-- -- 3 daysCephalic and trunk discs Paired trunk discs invaginate (Fig. 4) Pileus stageLarva develops transient lobes and lappets, the gut curves backward, the paired three days cerebral organ discs invaginate in the gut, and also the proboscis and dorsal rudiment seem (Figs. 3f and 5). Ciliary bands seem as four segments which span each and every transient lobe and lappet (Fig. six) The head and trunk rudiments fuse about the base in the gut (Fig. 8). Ciliary band segments are re-arranged to kind two full transverse ciliary bands Epidermis of trunk rudiment extends over the proboscis, but has not but fused with the epidermis of the head rudiment, leaving a dorsal gap (Fig. 9) The head and trunk rudiments fused to form a total juvenile (Fig. ten). Juvenile erupts from and devours larval physique in catastrophic metamorphosis 4 days 6 days 9 daysTorus stage Hood stage Metamorphosis-- -- 18 daysDash represents missing databands from the anterior and posterior lobes (Fig. 6b), too as anterior and posterior gaps separating the ciliary bands from the two lappets (Fig. 6a). The ciliary bands spanning every single from the transient lappets extend from a tiny pair of epidermal invaginations involving the anterior lobe and every single lappet (Fig. 5a, c and d, Extra file 2 -- Movie two, Added file three -- Film three). These "pits" are inside a position corresponding towards the anterior axils (the development zones) of the pilidium larva, which give rise for the ciliary bands in common pilidia [19]. At this point, the cephalic discs are positioned above the transient anterior lobe, when the trunk discs are beneath the backward curve with the gut, along the posterior side of every single lappet (Fig. 5b-d). In the same time, the unpaired dorsal rudiment types along the inner pilidial epidermis dorsal/apical to the gut, plus the unpaired proboscis rudiment types amongst the cephalic discs (Fig. 5). The origin of those rudiments is uncertain, but possibly, as is hypothesized for any typical pilidium, they may be mesenchymal [10]. At any price, they don't appear to kind by invagination from the larval epidermis. Moreover, two shallow chambers outpocket in the basal portion in the gut (close to the blastopore) among the building cephalic and trunk discs on either side of the mid-sagittal plane. These invaginations of your gut elongate and kind the cerebral organ discs (Fig. 5e3-f ). The larva also starts to develop musculature at this point. Initially, it develops circumferential muscles underlying the ciliary bands of your transitory larval lobes (which later form the anterior ciliary band). As these muscles type, some start to arc towards the posterior on the larva, extending into every single ofthe transitory lappets, instead of encircling the larva (Fig. 7a-b). These extensions type an "arc" of muscle which follows the curve on the lappet (consider two jump ropes, each and every wrapped around opposite sides on the larva, are permitted to droop into the lappet on its respective side).